Professor STEPHEN REDPATH
Chair in Conservation Science
Personal Details
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+44 (0)1224 274181 |
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Address: |
Aberdeen Centre for Environmental Sustainability (ACES), |
Biography
- 2008- Director of ACES
- 2007 Chair in Conservation Science, Aberdeen University.
- 1997-2007 Research Scientist at Centre for Ecology & Hydrology, Banchory.
- 1990-1997 Research Scientist at Institute of Terrestrial Ecology, Monks Wood.
- 1985-1989 PhD Leeds University.
- 1982-1985 BSc (Ecology) Leeds University.
Research Interests
My core research interests lie in animal ecology and conservation science. My work has focused on long-term and large-scale field systems, using experiments to tease out the impact of population processes and land use on individual behaviour, populations and communities. Within ACES we are linking natural, social and physical sciences together with policy makes and stakeholders to tackle key research questions in environmental sustainability.
My research group focuses on the following areas:
- Interacting processes and population dynamics
- Grazing and biodiversity
- Climate and migrating birds
- Predation and predator management
- Scaling in interdisciplinary research
Current Research
Interacting processes and population dynamics.
1) Red grouse, parasites & territorial behaviour. NERC funded. With Sasha Dall (UEC), Dan Haydon (Glasgow), Francois Mougeot & Stuart Piertney (Aberdeen), Jason Matthiopoulos (St. Andrews) and Pete Hudson (Penn State). One of the main challenges facing population ecology is to explain the enormous spatial and temporal variation observed in patterns of abundance. Our understanding of these patterns has been illuminated by detailed studies of species such as red grouse with unstable dynamics. Long-term harvest records of grouse show that populations exhibit unstable dynamics with geographical variation in cycle period ranging from 2 to 15 years. Within this system there are two dominant, destabilising biotic factors: territorial behaviour and parasites. We recently undertook a wide range of population level experiments that confirmed the de-stabilising role of both processes and explored how they interacted. Here, we seek to test the hypothesis that interactions between parasites, territorial behaviour and climate can account for the spatial variation in the dynamical behaviour of red grouse populations.
This project links in with the NERC Advanced Fellowship of Francois Mougeot - Endocrine-immune function trade-offs, genetic quality and honest sexual signalling. Francois is exploring the endocrine mechanisms that influence both sexual signals and immune function and how the genetic background of the individual moderates these effects. The grouse system therefore enables us to reach from genes to large-scale population dynamics.
2) From individual behaviour to population dynamics. As part of the NERC grant above (1), we will use game theory to explore the strategic consequences of the interaction between aggressive territorial behaviour and increased susceptibility to infection by parasites. In systems where multiple biotic factors are operating, individuals have to trade-off the risks and benefits of alternative strategies in order to maximise their fitness. The consequences of the decisions individuals take in these trade-offs may profoundly affect population size and dynamic stability. Moreover, given the trade-offs associated with territorial aggression, the fitness consequences of individual decisions are likely to depend on the decisions of others - for instance, the costs of fighting will depend on how aggressive opponents are likely to be. We will obtain specific parameters from this game theoretic approach and from new experiments to incorporate into a population model and compare the emergent dynamic properties against existing long-term time series.
3) Rabbits, predation risk & myxomatosis. 
Funded by Junta de communidades de Castilla La Mancha. With Rafael Villafuerte (UCLM). Rabbits are considered a keystone species in their native Spain. However, over the last 50 years, populations have undergone a sharp decline, mainly as a result of the arrival of two viral diseases, Myxomatosis in the 1950s and Rabbit Haemorrhagic Disease (RHD) at the end of the 1980s. Across Spain, rabbits are an important resource, being hunted in more than 30,000 private hunting estates, covering more than 70% of the country. The decline poses potentially profound consequences for hunting practices and for the wide guild of predators that is dependent on them, including the endangered Iberian Lynx (Lynx pardinus) and the Spanish Imperial Eagle (Aquila adalberti). There is a strong need to try and restore rabbit populations. Here, we seek to test the hypothesis that interactions between predation risk and disease prevent population recovery.
Grazing and biodiversity.
In 2002 we started a grazing manipulation in the Scottish uplands, to examine the consequences of changing herbivory on vegetation dynamics and the cascading effects through upland ecosystems. This work is funded by SEERAD and is led by Robin Pakeman (Macaulay Institute) and is in collaboration with Darren Evans (Bristol). Since 2002, we have focused on developing a mechanistic understanding of how grazing affects upland birds, with a focus on meadow pipits as a model system. We have found rapid effects of our manipulations on pipit egg size and pipit abundance. We have also found rapid effects on the abundance of field voles. Whilst continuing the long-term monitoring of vertebrate responses, we have two studentships (Nacho Villar & Gina Prior) working on this system.
4) Grazing, voles and predation - consequences of land use change in the uplands. Student - Nacho Villar. Other supervisors - Lambin (Aberdeen), Pakeman (Macaulay), Evans (Bristol). To optimize biodiversity in the uplands, the challenge is to find how to balance conservation objectives with other users. Grazing can profoundly affect biodiversity, either directly or indirectly. One important indirect route is through its effect on predators, with knock-on implications for upland ecosystems. The effects of grazing will vary with predator type, with high grazing pressure expected to lead to increased corvid abundance, but decreased field vole Microtus agrestis abundance and thus a decrease of a variety of other avian and mammalian predators. Nacho is exploring the interactions between grazing, voles and predation, and the implications for ecosystem structure and land use in the uplands.
5) Grazing, climate, and breeding success in upland birds. Student - Gina Prior. Other supervisors - Thirgood (Macaulay), Monaghan (Glasgow), Evans (Bristol). Few studies have developed a mechanistic understanding of how grazing and climate affect upland species, and equally importantly, how they interact to influence their viability. This studentship will investigate the consequences to birds of varying climate (using altitude as a surrogate) and grazing in the uplands. It will use an upland passerine, the meadow pipit, as a model species, being an abundant, easy to study insectivore and a good bio-indicator of ecosystem health. The studentship will explore the mechanisms linking changes in climate and grazing with egg size and breeding success and how this subsequently links with density. Gina is exploring the plausible mechanisms linking climate and grazing to bird abundance. The experimental design will allow her to examine how the effects of climate change on birds may be modified by altering grazing regimes.
Climate and migrating birds.
6) Migration decisions in a changing world - mechanisms and consequences. Student - Alexandra Komissarova. Other supervisors - Travis (Aberdeen), Dawson (CEH).There is increasing evidence that many species are adapting to climate change by altering migration strategies. However, the mechanisms and consequences of such changes are poorly understood. A common strategy among a variety of taxa, including 60% of European birds, is partial migration, where some individuals within populations over-winter on breeding sites, whilst others leave. This strategy presents an ideal model for developing a mechanistic understanding of migration as it allows for comparison at an individual level, without the problems associated with cross-species analyses. Alex is comparing strategies across an environmental gradient and using an experimental approach to test the key hypothesis that the probability of migrating is dependent on condition and frequency-dependent effects (eg social rank), but that the threshold for migration will vary between sites. She works the robin, a relatively well-studied partial migrant, in north east Scotland and in England.
7) Modelling the impact of climate change on migratory birds. Student - Karen Mustin. Other supervisors - Travis (Aberdeen), Sutherland (Cambridge), Gill (UEA), Thomas (York). While all species are faced with a changing climate, for return migrants, the challenge is potentially the greatest. In this project, Karen is developing a combination of statistical and individual-based models to improve our understanding of the likely impacts of climate change on migratory birds. The work includes a mix of strategic models designed to develop generic insight into ecological and evolutionary dynamics of migratory species' ranges, and tactical modelling aimed at predicting the future ranges of selected bird species that have either their summer or winter ranges in Britain.
8) Diagnosing the causes of Ring Ouzel declines in Britain. Student - Innes Sim. Other supervisors - Reid (Aberdeen).The Ring Ouzel is a summer migrant to Britain, where it is associated with upland areas, in particular steep sided valleys, crags and gullies. There has been concern for some time that the UK Ring Ouzel population is in decline and in 2002, Ring Ouzel was placed on the Red List in Birds of Conservation Concern, due to an estimated 58% population decline between 1988-91 and 1999. The underlying causes of this serious decline are unknown, and may be linked to problems in breeding or wintering areas, or on migration routes. However, a recent correlative analysis suggests that declines were more likely in years following high, late-summer temperatures. This suggests direct or indirect climate effects on late breeding attempts and/or post fledging and pre-migratory body condition, probably through a reduction in food availability. Innes will be exploring how moorland ecosystems and climate interact to influence breeding site occupancy, productivity, chick condition and subsequent post-fledging survival to provide future management prescriptions aimed at reversing Ring Ouzel declines.
Predation and predator management.
9) Hen harriers - prediction, impact & mitigation. Part funded by UKPopNET. Collaborators - Anderson (York), Travis (Aberdeen). One of the most contentious conflicts in UK conservation has been over the management of Hen Harriers on grouse moors. We have worked on this problem since 1986, quantifying the impact of harriers on grouse populations through long-term studies and exploring mitigation strategies. The work is currently continuing in two ways. Firstly, we are building static and dynamic spatial models to predict harrier distributions across the UK, highlight conflict areas and testing alternative management strategies. Second, we are involved in the ongoing stakeholder dialogue, run by the Environment Council with the aim of exploring ways forward to resolve the conflict.
10) Rabbits, predators and the predator-pit hypothesis. Funded by Spanish Government, in collaboration with Ferreras and Villafuerte (UCLM). Here, we are exploring the role of predation in explaining the lack of recovery of wild rabbit populations in centre-southern Iberian Peninsula, after the arrival of the rabbit haemorrhagic disease (RHV) at the end of the 1980’s. According to the predator pit hypothesis, predation would have a regulatory effect at low rabbit density, preventing a return to the initial, high-density equilibrium. We have previously shown that most rabbit populations in the centre-southern Iberian Peninsula remained low after the arrival of the RHD. However, so far no study has assessed whether this lack of recovery in rabbit populations is related to a predation-mediated equilibrium. In this project our objectives are: 1. To assess the possibility that rabbit populations are regulated at low density by predation in the centre-southern Iberian Peninsula. 2. To quantify the joint impact of the predator community on rabbits, comparing localities with similar habitat characteristics, where rabbits are at different densities. 3. To assess functional and numerical responses of predators to changes in the abundance of rabbit populations. and 4. To propose management strategies which enable the effective recovery of rabbit populations.
Scaling in interdisciplinary research.
11) Scaling in interdisciplinary upland research: from plots to catchments to regions. Funded by UKPopNET. Led by Travis (Aberdeen) in collaboration with Dytham (York). PDRA - Steve Palmer. Due to start August 2007. Scaling is an important part of the problem solving process for many disciplines; ecologists, biogeochemists, sociologists and economists all deal with scale-dependent issues. Frequently we scale-down to gain a mechanistic understanding of processes before using models to upscale and make predictions. Only very infrequently have we properly validated this method and this has led to considerable problems in persuading policy makers of the usefulness of plot-scale research. We propose to take advantage of the unique opportunity provided by the large-scale UKPopNET upland experiment by taking an interdisciplinary approach to test and develop scaling approaches including the use of spatial statistics, multi-level statistics, and GIS-based modelling.
Collaborations
I collaborate widely with a range of colleagues locally, nationally and internationally.
Within ACES my colleagues are: Ben Davies, Antonio Ioris, Eva Krupp, Andy Taylor, Simon Thirgood and Rene Van der Wal.
My other principal collaborators:
Barb Anderson (York University); Beatriz Arroyo (IREC); Sasha Dall (University Exeter in Cornwall); Alastair Dawson (CEH); Darren Evans (Bristol); Pablo Ferreras (IREC); Dan Haydon (Glasgow); Peter Hudson (Penn State); Xavier Lambin (Aberdeen), Jason Matthiopoulos (St. Andrews)Francois Mougeot (Aberdeen & CSIC); Robin Pakeman (Macaulay); Stuart Piertney (Aberdeen); Jane Reid (Aberdeen); Justin Travis (Aberdeen); Rafael Villafuerte (IREC).
Research Grants
CURRENT:
2008-2011 EU FP7. Hunting for sustainability. With Thirgood (Macaulay) and partners across Europe & Africa.
2007-2010 NERC. The impact of interacting processes on population dynamics. With Dall (UEC) & Haydon (Glasgow).
2007-2009 UKPopNET. Scaling in interdisciplinary research: from plots to catchments to regions. With Travis - PI (Aberdeen) and Pitchford (York).
2006-2008 Spanish Government. Relationships between predation risk and disease in the wild rabbit. With Villafuerte - PI & Ferrera (CSIC).
2006-2008 Spanish Government. Quantifying the predator pit hypothesis: predation and the recovery of wild rabbit populations in Spain. With Ferrera - PI & Villafuerte (CSIC).
RECENT:
2004-2006 UKPopNET NERC. A framework for sustainable livelihoods, biodiversity change and conflict resolution.With White (CEH), Raffaelli (York), Watkinson (UEA), Bateman (UEA), Lambin (Aberdeen) & Richards (MI).
2002-2006. SNH. Habitat use and range management of priority areas for hen harriers. With Arroyo (CEH).
2000-2004 NERC. Red grouse population cycles: territorial behaviour and parasites. With Hudson (Penn State).
External Responsibilities
Member of NERC Peer Review Panel.
Member of Moorland Forum.
Board member of Heather Trust.
Member of Science Management Committee of UKPopNET.
Editor for Animal Conservation.
PhD Students
Current:
Innes Sim (start 2007) Diagnosing the causes of Ring Ouzel declines in Britain. RSPB funded. Co-supervisors: Reid.
Gina Prior (start 2006). Climate change, grazing and breeding success in upland birds. MDT funded. Co-supervisors: Thirgood, Monaghan & Evans.
Alexandra Komissarova (start 2006) Migration decisions in a changing world - mechanisms and consequences . NERC funded. Co-supervisors: Travis & Dawson
Karen Mustin (start 2006) Modelling the impact of climate change on migratory birds. UKPopNET funded. Co-supervisors: Travis.
Nacho Villar – (start 2005). Grazing, voles and predation – consequences of land use change in the uplands. MI/ACES funded. Co-supervisors: Lambin, Pakeman & Evans.
Graduated:
Graham Rebecca (2006). Impact of commercial forestry on the population ecology of Merlins in north-east Scotland.
Kerry Lock (2004). Role of nepotism in driving red grouse population cycles.
Linzi Seivwright (2004). Patterns of transmission of T. tenuis in Red Grouse.
Arjun Amar (2001). Conservation of declining hen harriers Circus cyaneus in Orkney: roles of land use and polygyny.
Selected Publications
Some Recent Publications
Thirgood, S.J. & Redpath, S.M. (2008). Hen Harriers and Red Grouse: science, politics and human-wildlife conflict. Journal of Applied Ecology - In Press.
Piertney, S.P, Lambin, X., Maccoll, A.D., Lock, K., Bacon, P.J., Dallas, J.F., Leckie, F., Mougeot, F., Racey, P.A., Redpah, S. & Moss R. (2008) Temporal changes in kin structure through a population cycle in a territorial bird, the red grouse Lagopus lagopus scoticus. Molecular Ecology 17: 2544-2551.
Mougeot, F., Perez-Rodriguez, L., Martinez-Padilla, J. & Redpath, S.M. (2007) Parasites, testosterone and honest carotenoid-based signalling of health. Functional Ecology 21: 886-898.
Delibes-Mateos, M., Redpath, S.M., Angulo, E., Ferreras, P. & Villafuerte, R. (2007) Rabbits as a keystone species in southern Europe. Biological Conservation. 137: 149-156.
Evans, D.M., Redpath, S.M., Evans, S.A., Elston, D.A., Gardner, C., Dennis, P. & Pakeman, R. (2006) Low intensity, mixed livestock grazing improves the breeding abundance of a common insectivorous passerine. Biology Letters 2: 636-638.
Redpath, S.M., Mougeot, F., Leckie, F.M. & Hudson, P.J. (2006) Testing the role of parasites in driving the cyclic population dynamics of a gamebird. Ecology Letters 9:410-418.
Redpath, S.M., Mougeot, F. & Leckie, F.M. (2006) The effects of autumn testosterone on survival and productivity in red grouse Lagopus lagopus scoticus. Animal Behaviour 71: 1297-1305
Seivwright, L., Redpath, S.M., Mougeot, F. & Hudson, P.J. (2005) Interactions between intrinsic and extrinsic mechanisms in red grouse (Lagopus lagopus scoticus). Proc. Roy. Soc.B. 272: 2299-2304.
Mougeot, F., Redpath, S.M. & Piertney, S.B. (2005) Teasing apart behavioural and physiological mechanisms in testosterone mediated trade-offs. American Naturalist 166 (2): 158-168.
Mougeot, F., Piertney, S. B., Leckie, F., Evans, S., Moss, R., Redpath, S. & Hudson, P. J. (2005) Experimentally increased aggressiveness reduces population kin structure and subsequent recruitment in red grouse Lagopus lagopus scoticus. Journal of Animal Ecology 74: 488-498.
Redpath, S.M., Arroyo, B.E., Leckie, F,M., Bacon, P., Bayfield, N., Gutiérrez, R.J. & Thirgood, S.J. (2004)Using decision modelling with stakeholders to reduce human-wildlife conflict: a raptor - grouse case study. Conservation Biology: 18, 350-359.
Mougeot, F., Redpath, S.M., Hudson, P.J., Moss, R. & Matthiopolous J. (2003) Territorial behaviour and population dynamics in red grouse Lagopus lagopus scoticus. I. Population level experiments. J. Anim. Ecol. 72: 1073-1082.
Mougeot, F. Redpath, S.M. Leckie F. & Hudson P.J.(2003) The effect of aggressiveness on the population dynamics of a territorial bird. Nature 421: 737-739.
Full Publication List.
White R, Fischer A, Marshall K, Travis J, Webb T, Di Falco S, Redpath S & Van der Wal R (2008). Integration of social, economic and ecological perspectives to resolve biodiversity conflicts: development of a conceptual framework. Land Use Policy - in press.
Van der Wal R, Fischer A, MarquissM, RedpathS & Wanless S (2008). Is bigger necessarily better for environmental research? Scientometrics - in press.
Martinez, J.E., Martinez, J.A., Zuberogoitia, I., Zabala, J., Redpath, S.M. & Calvo, J.F. (2008). The effect of intra- and interspecific interactions in the large sclae disctribution of cliff nesting raptors. Ornis Fennice - In Press.
Baines, D., Redpath., S., Richardosn, M. & Thirgood, S. (2008). The direct and indirect effects of predation by Hen Harriers on trends in breeding birds on a Scottish grouse moor. Ibis - In Press.
Leckie, F.M., Arroyo, B.E., Thirgood, S.J. & Redpath, S.M. (2008). Parental differences in brood provisioning by Hen Harriers Circus cyaneus. Bird Study - In Press.
Thirgood, S.J. & Redpath, S.M. (2008). Hen Harriers and Red Grouse: science, politics and human-wildlife conflict. Journal of Applied Ecology - In Press.
Amar, A., Thirgood, S, Pearce-Higgins, J. & Redpath, S.M. (2008). The impact of raptors on the abundance of upland passerines and waders. Oikos - In Press. doi: 10.1111/j.2008.0030-1299.16769.x,
Piertney, S.P, Lambin, X., Maccoll, A.D., Lock, K., Bacon, P.J., Dallas, J.F., Leckie, F., Mougeot, F., Racey, P.A., Redpah, S. & Moss R. (2008) Temporal changes in kin structure throgh a population cycle ina territorial bird, the red grouse Lagopus lagopus scoticus. Molecular Ecology 17: 2544-2551.
Martinez, J, A., Calvo, J.F., Martinez, J.E., Zuberogoitia, I., Zabal, J. & Redpath, S.M. (2008). Breeding performance, age effects and territory occupancy in a Bonelli's Eagle Hieraaetus fasciatus population. Ibis 150: 223-233.
Amar, A., Arroyo, B., Meek, B., Redpath, S.M. & Riley, H. (2008) Influence of habitat on occupancy and breeding performance of hen harriers Circus cyaneus in Orkney. Ibis 150: 400-404
Mougeot, F., Perez-Rodriguez, L., Martinez-Padilla, J. & Redpath, S.M. (2007) Parasites, testosterone and honest carotenoid-based signalling of health. Functional Ecology 21: 886-898.
Irvine, R.J., Leckie, F.M. & Redpath, S.M. (2007). The cost of carrying radio-transmitters: a test with homing pigeons. Wildlife Biology 13 (3): 238-243
Matthiopoulos, J., Graham, K., Smout, S., Asseburg, C., Redpath, S.M., Thirgood S.J., Hudson, P. & Harwood, J. (2007) Sensitivity to assumptions in models of generalist predation on a cyclic prey. Ecology 88(10): 2576-2586.
Delibes-Mateos, M., Redpath, S.M., Angulo, E., Ferreras, P. & Villafuerte, R. (2007) Rabbits as a keystone species in southern Europe. Biological Conservation. 137: 149-156.
Mougeot, F. & Redpath, S.M. (2007) Testosterone and parasite-mediated sexual selection in a tetraonid bird. In: Testosterone Research Trends. Ed L.I.Ardis, Nova Science Publishers, New York.
Evans, S.A., Mougeot, F., Redpath, S.M. & Leckie, F. (2007) Alternative methods for estimating breeding density in an upland game bird, the red grouse Lagopus lagopus scoticus. Wildlife Biology. 13(2): 130-139.
Ellis, D. H. Lish, J. W. Kéry, M. & Redpath, S. M. (2007) Short term oscillations in avian molt intensity: evidence from the Golden Eagle Aquila chrysaetos. J. Avian. Biol. 37: 642-644.
Evans, D.M., Redpath, S.M., Evans, S.A., Elston, D.A., Gardner, C., Dennis, P. & Pakeman, R. (2006) Low intensity, mixed livestock grazing improves the breeding abundance of a common insectivorous passerine. Biology Letters 2: 636-638.
Redpath, S.M., Mougeot, F., Leckie, F.M. & Hudson, P.J. (2006) Testing the role of parasites in driving the cyclic population dynamics of a gamebird. Ecology Letters 9:410-418.
Redpath, S.M., Leckie, F.M., Thirgood, S.J. & Amar, A. (2006). Compensating for the costs of polygyny in hen harriers Circus cyaneus. Behav. Ecol & Sociobiol. 60 (3): 386-391
Redpath, S.M., Mougeot, F. & Leckie, F.M. (2006) The effects of autumn testosterone on survival and productivity in red grouse Lagopus lagopus scoticus. Animal Behaviour 71: 1297-1305
Evans, DM, Redpath, SM, Elston, DA, Evans, SA & Mitchell, RJ. (2006) To graze or not to graze? Sheep, voles, forestry and nature conservation in the British uplands. J. Appl Ecol. 43: 499-505.
Sunde, P. & Redpath, SM (2006) Measuring spatial responses to habitat heterogeneity: owls in fragmented forest. Ecography29: 152-158.
Mougeot, F, Redpath, SM & Piertney, S.B. (2006) Elevated spring testosterone increases parasite intensity in male red grouse. Behavioral Ecology 17:117-125.
Seivwright, L., Redpath, S.M., Mougeot, F. & Hudson, P.J. (2005) Interactions between intrinsic and extrinsic mechanisms in red grouse (Lagopus lagopus scoticus). Proc. Roy. Soc.B. 272: 2299-2304.
Evans, D.M., Redpath, S.M. & Evans, S.A. (2005) Seasonal patterns in the productivity of Meadow Pipits in the uplands of Scotland. J. Field Orn.76: 245-251.
Mougeot, F., Dawson, A., Redpath, S. & Leckie, F. (2005)Testosterone and autumn territorial behaviour in male red grouse Lagopus Lagopus Scoticus. Hormones & Behavior 47:576-584.
Mougeot, F., Redpath, S.M. & Leckie, F.M. (2005) Ultra-violet reflectance of male and female red grouse (Lagopus lagopus scoticus) sexual ornaments reflect nematode parasite intensity.J. Avian Biology 36(3): 203-209.
Mougeot, F., Redpath, S.M. & Piertney, S.B. (2005) Teasing apart behavioural and physiological mechanisms in testosterone mediated trade-offs. American Naturalist 166 (2): 158-168.
Mougeot, F., Evans, S.A. & Redpath, S.M. (2005). Interactions between population processes in a cyclic species: parasites reduce autumn territorial behaviour of male red grouse Lagopus lagopus scoticus. Oecologia 144 (2): 289-298.
Mougeot, F., Piertney, S. B., Leckie, F., Evans, S., Moss, R., Redpath, S. & Hudson, P. J. (2005) Experimentally increased aggressiveness reduces population kin structure and subsequent recruitment in red grouse Lagopus lagopus scoticus. Journal of AnimalEcology 74: 488-498.
Evans, D.M., Redpath, S.M., Evans, S.A. & Elston, D.A. (2005) Livestock grazing effects the egg size of an insectivorous passerine. Biology Letters 1(3):322-325.
Thirgood S.J.& Redpath S.M. (2005). Science, politics and human-wildlife conflicts: harriers and grouse in the UK. In: People or Wildlife: Conflict or Coexistence. Eds Woodroffe, R., Thirgood, S. & Rabinowitz, A. ZSL. London.
Valkama.J., Korpimäki.E., Arroyo.B., Beja. P, Bretagnolle. V., Bro.E., Kenward. R., Mañosa. S., Redpath.S.M., Thirgood. S.J., and Viñuela.J. (2005). Birds of prey as limiting factors of gamebird populations in Europe: a review. Biological Reviews 80:171-203
Mougeot F., Irvine J., Seivwright L., S.M Redpath & S. Piertney. (2004) Testosterone, immunocompetence and honest sexual signalling in male red grouse. Behavioral Ecology15:930-937.
Amar, A., Arroyo, B., Redpath, S.M. & Thirgood, S.J. (2004)Habitat predicts losses of red grouse to individual hen harriers. J. Appl. Ecol. 41: 305-314.
Amar, A., Picozzi, N, Meek, E.R., Lambin, X. & Redpath, S.M. (2004). The decline of the Hen Harrier on Orkney: exploring the changes to demographic parameters and mating system.Bird study52:18-24.
Amar, A. & Redpath, S.M. (2004). The implications of land use change on the decline of a top predator: the Hen Harrier on the Orkney Islands, Scotland. Ibis147:37-47.
Mougeot, F. & Redpath, S.M. (2004). Sexual ornamentation relates to immune function in male red grouse Lagopus lagopus scoticus. J. Avian Biol.35: 425-433.
Seivwright, L. J. Mougeot, F. Redpath, S. M. Watt, L. & Hudson P. J. (2004)Measuring nematode parasite burdens in red grouse (Lagopus lagopus scoticus): a methodological review. Jnl. Helminthology 78: 69-76.
Redpath, S.M., Arroyo, B.E., Leckie, F,M., Bacon, P., Bayfield, N., Gutiérrez, R.J. & Thirgood, S.J. (2004)Using decision modelling with stakeholders to reduce human-wildlife conflict: a raptor - grouse case study. Conservation Biology: 18, 350-359.
Redpath, S.M. & Appleby, B. (2004) The hooting of Tawny Owls: Structure and Function. In Nature's music: the science of birdsong. Eds.H Slabbekoorn & P. Marler. Academic Press, New York.
Matthiopolous J., Moss, R., Mougeot, F., Lambin, X. & Redpath, S.M. (2003) Territorial behaviour and population dynamics in red grouse Lagopus lagopus scoticus. II. Population models. J. Anim. Ecol. 72: 1083-1096.
Mougeot, F., Redpath, S.M., Hudson, P.J., Moss, R. & Matthiopolous J. (2003) Territorial behaviour and population dynamics in red grouse Lagopus lagopus scoticus. I. Population level experiments. J. Anim. Ecol. 72: 1073-1082.
Mougeot, F. Redpath, S.M. Leckie F. & Hudson P.J.(2003) The effect of aggressiveness on the population dynamics of a territorial bird. Nature 421: 737-739.
Amar, A, Redpath, S.M. & Thirgood, S.J. (2003) Evidence for food limitation in the declining hen harrier population on the Orkney islands, Scotland. Biological Conservation 11: 377-384.
Amar, A., Redpath, S., Lambin X. & Meek, E. (2003) Could the Hen Harrier (Circus cyaneus) decline on Orkney be due to a shortage of food? In: Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh. Thompson, DBA, Redpath, SM, Fielding AH, Marquiss, M. & Galbraith CA (Eds).
Redpath, S.M. & Thirgood, S.J. (2003)The impact of Hen Harrier (Circus cyaneus) predation on Red Grouse (Lagopus l. scoticus) populations: linking models with field data. Pages 499-511. In: Birds of Prey in a Changing Environment. The Stationery Office, Edinburgh. Thompson, DBA, Redpath, SM, Fielding AH, Marquiss, M. & Galbraith CA (Eds).
Redpath, S.M., Arroyo, B.E., Etheridge, B, Leckie, F.M., Bouwman, K. & Thirgood, S.J. (2002) Weather and harrier productivity: from local mechanisms to geographical patterns. Ecography 25: 533-540.
Thirgood, S.J., Redpath, S.M., Campbell, S. & Smith, A. (2002). Do habitat characteristics influence predation on red grouse? J. Appl. Ecol.39: 217-225.
Thirgood, S.J., Redpath, S.M. and Graham, I. (2002). What determines the foraging distribution of raptors on heather moorland. Oikos 100: 15-24.
Redpath, S.M., Amar, A., Madders, M. Leckie, F.M. & Thirgood, S.J. (2002) Hen harrier foraging success in relation to land use in Scotland. Anim. Cons. 5: 113-118.
Amar, A & Redpath, S.M. (2002) Determining the cause of the hen harrier decline on the Orkney Islands: an experimental test of two hypotheses. Anim. Cons. 5: 21-28.
Redpath, S.M., Thirgood, S.J. & Clarke, R.G. (2002) Field Vole abundance and Hen Harrier diet and breeding success in Scotland. Ibis. 144: E33-E39.
Campbell, S., Smith, A., Thirgood, S. & Redpath, S. (2002) Nest site characteristics and nesting success in red grouse Lagopus lagopus scoticus. Wildlife Biology. 8: 169-175.
Redpath, S.M., Thirgood, S.J. & Leckie, F. (2001) Does supplementary feeding reduce predation of Red Grouse by Hen Harriers? J. Appl. Ecol. 38: 1157-1168.
Smith, A., Redpath, S.M., Thirgood, S.J. & Campbell, S. (2001)Habitat characteristics of managed grouse moors and the abundance of meadow pipits and red grouse. J. Appl. Ecol. 38: 390-401.
Redpath, S.M., Clarke, R.G., Madders, M. & Thirgood, S.J. (2001). Assessing raptor diet: comparing pellets, prey remains and observational data at hen harrier nests. The Condor. 103: 184-188.
Redpath, S.M., Appleby, B.M. & Petty, S.J. (2000) Honest signalling in Tawny Owls – Do male hoots betray parasite loads. J. Avian Biol. 31: 457-462.
Smith, A., Redpath, S.M. & Campbell, S. (2000) The influence of moorland management on grouse and their predators. Final Report to the DETR. HMSO publication, The Stationery Office.
Thirgood, S.J., Redpath, S.M., Newton, I., Rothery, P. & Aebischer, N. (2000). Raptor predation and population limitation in red grouse. J. Anim. Ecol. 69: 504-516.
Thirgood, S.J., Redpath, S.M., Newton, I. & Hudson, P. (2000). Raptors and grouse: conservation conflicts and management solutions. Cons. Biol. 14: 95-104.
Thirgood, S.J. Redpath, S.M., Haydon, D.T., Rothery, P.,Newton, I. & Hudson, P.J. (2000) Habitat loss and raptor predation: disentangling long- and short-term causes of red grouse declines. Proc. R. Soc. Lond. B. 267: 651-656.
Redpath, S.M. & Thirgood, S.J. (1999) Functional and numerical responses in generalist predators: hen harriers and peregrines on Scottish grouse moors. J. Anim. Ecol. 68: 879-892.
Leckie, F., May, R., Thirgood, S.J., Donnelly, E. & Redpath, S.M. (1998) The diet of foxes on moorland managed for red grouse. Journal of Zoology. 21: 599-604.
Thirgood, S.J. & Redpath, S.M. (1997) Red grouse and their predators. Nature 390: 547.
Thirgood, S.J., Redpath, S.M. & Hudson, P.J. (1997) Estimating the cause and rate of mortality in red grouse Lagopus lagopus scoticus. Wildlife Biology 4: 65-71
Appleby, B.M. & Redpath, S.M. (1997) Geographical variation in the hoot of the tawny owl. Ibis 139: 152-158.
Appleby, B.M. & Redpath, S.M. (1997) Indicators of male quality in the hoots of tawny owls Strix aluco. J. Rapt. Res. 31: 65-70.
Thompson, D.B.A., Gillings, S.D., Galbraith, C.A., Redpath, S.M. & Drewitt, J. (1997). The contribution of game management to biodiversity: a review of the importance of grouse moors for upland birds. In: Biodiversity in Scotland. Eds. Fleming, Newton, Vickery & Usher. The Stationery Office, Edinburgh.
Redpath, S.M., Madders, M., Donnelly, E., Anderson, B., Thirgood, S.J. & McLead, D. (1997) Nest site selection by hen harriers on grouse moors. Bird Study 45: 51-61.
Redpath, S.M. & Thirgood, S.J. (1997) Birds of prey and red grouse. The final report of the Joint Raptor Study. HMSO publication.
Galeotti, P.R., Appleby, B.M. & Redpath, S.M. (1996) Macro and microgeographical variations in the 'hoot' of Italian and English tawny owls Strix aluco. Ital. J. Zool. 63: 57-64.
Redpath, S.M. (1995) Habitat fragmentation and the individual: tawny owls Strix aluco in woodland patches. J. Anim. Ecol. 64: 652-661.
Redpath, S.M. (1995) Impact of habitat fragmentation on activity and hunting behaviour in the tawny owl Strix aluco. Behav Ecol. 6: 410-415.
Thirgood, S.J., Redpath, S.M., Hudson, P.J., Hurley, M & Aebischer, N.J. (1995). Effect of necklace radio transmitters on survival and breeding success of red grouse. Wildlife Biology 1: 121-126.
Redpath, C.J., Thirgood, S.J. & Redpath, S.M. (1995) Evaluation of methods to estimate vole abundance in upland habitats. Journal of Zoology, London 237: 49-55.
Thirgood, S.J., Leckie, F.M. & Redpath, S.M. (1995) Diurnal and seasonal variation in line transect counts of moorland passerines. Bird Study 42: 257-259.
Graham, I.M., Redpath, S.M. & Thirgood, S.J. (1995) The diet and breeding density of Common Buzzards in relation to prey abundance. Bird Study 42: 165-173.
Redpath, S.M. (1994) Censusing tawny owls Strix aluco using imitation calls. Bird Study 41: 192-198.
Redpath, S.M. & Wyllie, I. (1994) Traps to capture territorial owls. J. Rapt. Res. 28: 115-117.
Redpath, S.M. (1992). Behavioural interactions between Hen Harriers and their moorland prey. Ornis Scandinavica 23: 73-80.
Redpath, S.M. (1991). The effects of hen harrier predation on red grouse breeding success. Journal of Applied Ecology. 28: 659-671.
Redpath, S.M. (1990). Hen harriers facilitating predation of red grouse deaths by buzzards. British Birds, 83(3): 116.
Reynolds, J.C., Angelstam, P. & Redpath, S.M. (1988). Predators, their ecology and impact on gamebird populations. In: Ecology and Management of gamebirds. Eds. P.J. Hudson and M.R.W. Rands. Blackwell Scientific Press, Oxford.
Redpath, S.M. (1988). Vigilance levels in preening dunlin (Calidris alpina). Ibis 130(4): 55-57.